copyright Rafał Zwolak



Zwolak R., P. Celebias, M. Zduniak, M. Bogdziewicz and A. Wróbel. 2024. Scatterhoarder abundance and advantages of seed burial drive dynamics of a tree-rodent interaction. Journal of Ecology. PDF

Zwolak R., D. Clement, A. Sih, and S. J. Schreiber. 2024. Granivore abundance shapes mutualism quality in plant-scatterhoarder interactions. New Phytologist, 241: 1840-1850. PDF

Celebias P., A. Wróbel, M. Zduniak, M. Steele, M. Bogdziewicz, and R. Zwolak. 2024. Behaviour, body mass and sex: How intraspecific variation influences seed dispersal by a scatter-hoarding rodent. Functional Ecology, 38: 1420-1430. PDF

Bogdziewicz M., (...), R. Zwolak, M. Żywiec, and A. J. Hacket-Pain. 2024. Evolutionary ecology of masting: mechanisms, models, and climate change. Trends in Ecology and Evolution.


Zduniak M., S. Serafini, A. Wróbel, and R. Zwolak. 2023. Host's body mass, not sex, affects ectoparasite loads in yellow-necked mouse Apodemus flavicollis. Parasitology Research, 122: 2599-2607. PDF

Baláž I., M. Bogdziewicz, S. Dziemian-Zwolak, C. Lo Presti, A. Wróbel, M. Zduniak, and R. Zwolak. 2023. From fleas to trees: masting indirectly affects flea abundance on a rodent host. Integrative Zoology, 18: 440-452. PDF


Zwolak R., P. Celebias, and M. Bogdziewicz. 2022. Global patterns in the predator satiation effect of masting - a meta-analysis. Proceedings of the National Academy of Sciences, 119: e2105655119. PDF

Bogdziewicz M., B. Jędrzejewska, E. Wysocka-Fijorek, A. Gazda, S. Miścicki, D. Kuijper, R. Zwolak, M. Churski, and T. Podgórski. 2022. Emerging infectious disease triggered a trophic cascade and enhanced recruitment of a masting tree. Proceedings of the Royal Society B, 289: 20212636. PDF

Bednarz P. and R. Zwolak. 2022. Body mass and sex, but not breeding condition and season, influence open-field exploration in the yellow-necked mouse. Ecology and Evolution 12: e8771. PDF

Wróbel A., P. Kurek, M. Bogdziewicz, D. Dobrowolska, and R. Zwolak. 2022. Avian dispersal of an invasive oak is modulated by acorn traits and the presence of a native oak. Forest Ecology and Management 505: 119866. PDF


Zwolak R., D. Clement, A. Sih, and S. J. Schreiber. 2021. Mast seeding promotes evolution of scatter-hoarding. Philosophical Transactions of the Royal Society B, 376: 202003754. PDF


Zwolak R., M. Bogdziewicz M., and E. E. Crone. 2020. On the need to evaluate costs and benefits of synzoochory for plant populations. Journal of Ecology, 108: 1784-1788. PDF

Zwolak R., and A. Sih. 2020. Animal personalities and seed dispersal: A conceptual review. Functional Ecology, 34: 1294-1310. PDF

Bogdziewicz M., E. E. Crone, and R. Zwolak. 2020. Do benefits of seed dispersal and caching by scatterhoarders outweigh the costs of predation? An example with oaks and yellow-necked mice. Journal of Ecology, 108:1009-1018. PDF


Schupp E. W., R. Zwolak (...), and K. Shea (14 authors). 2019. Intrinsic and extrinsic drivers of intraspecific variation in seed dispersal are diverse and pervasive. AoB Plants 11: plz067. PDF

Meagher S., K. L. Winters, K. W. McCravy, and R. Zwolak. 2019. Complex and diverse drivers of parasite loads in a cosmopolitan insect. Journal of Parasitology 105: 659-668. PDF

Wróbel A., E. E. Crone, and R. Zwolak. 2019. Differential impacts of soil microbes on native and co-occurring invasive tree species. Ecosphere 10: e02802.  PDF

Snell R. S., N. G. Beckman, (...), R. Zwolak, and E. Schupp (22 authors). 2019. Consequences of intraspecific variation in seed dispersal for plant demography, communities, evolution, and global change. AoB Plants 11: plz016. PDF

Wróbel A. and R. Zwolak. 2019. Habitat-dependent seed dispersal of an introduced tree species by native rodents. Forest Ecology and Management 433: 563-568. PDF

Bogdziewicz M., N. I. Lichti, and R. Zwolak. 2019. Consumer-mediated indirect interaction with a native plant lowers the fitness of an invasive competitor. Journal of Ecology 107: 12-22. PDF


Zwolak R. 2018. How intraspecific variation in seed-dispering animals matters for plants. Biological Reviews 93: 897-913. PDF

Bogdziewicz M., S. Marino, R. Bonal, R. Zwolak, and M. A. Steele. 2018. Rapid aggregative and reproductive responses of weevils to masting of North American oaks counteract predator satiation. Ecology 99: 2575-2582. PDF

Zwolak R., J. Witczuk, M. Bogdziewicz M., L. Rychlik, and M. Pagacz. 2018. Simultaneous population fluctuations of rodents in montane forests and alpine meadows suggest indirect effects of tree masting. Journal of Mammalogy 99: 586-595. PDF

Bogdziewicz M., R. Bonal, J. M. Espelta, E. M. Kalemba, M. A. Steele, and R. Zwolak. 2018. Invasive oaks escape pre-dispersal insect seed predation and trap enemies in their seeds. Integrative Zoology 13: 228-237. PDF


Wróbel A. and R. Zwolak. 2017. Deciphering the effects of disperser assemblages and seed mass on patterns of seed dispersal in a rodent community. Integrative Zoology 12: 457-467. PDF

von Merten S., R. Zwolak, and L. Rychlik. 2017. Social personality: a more social shrew species exhibits stronger differences in personality types. Animal Behaviour 127: 125-134. PDF

Bogdziewicz M., E. E. Crone, M. A. Steele, and R. Zwolak. 2017. Effects of nitrogen deposition on reproduction in a masting tree: benefits of higher seed production are trumped by negative biotic interactions. Journal of Ecology 105: 310-320. PDF


Bogdziewicz M., R. Zwolak, L. Redosh, L. Rychlik, and E. E. Crone. 2016. Negative effects of density on space use of small mammals differ with the phase of the masting-induced population cycle. Ecology and Evolution 6:8423-8430. PDF

Zwolak R., M. Bogdziewicz, A. Wróbel, and E. E. Crone. 2016. Advantages of masting in European beech: timing of granivore satiation and benefits of seed caching support the predator dispersal hypothesis. Oecologia 180: 749-758. PDF

Bogdziewicz M., R. Zwolak, and E. E. Crone. 2016. How do vertebrates respond to mast seeding? Oikos 125: 300-307. PDF

Zwolak R., M. Bogdziewicz, and L. Rychlik. 2016. Beech masting modifies the response of rodents to forest management. Forest Ecology and Management 359: 268-276. PDF


Dziemian S., B. Sikora, B. Piłacińska, J. Michalik, and R. Zwolak. 2015. Ectoparasite loads in sympatric urban populations of the northern white-breasted and the European hedgehog. Parasitology Research 114: 2317-2323. PDF


Dziemian S., J. Michalik, B. Piłacińska, S. Bialik, B. Sikora, and R. Zwolak. 2014. Infestation of urban populations of the Northern white-breasted hedgehog, Erinaceus roumanicus, by Ixodes spp. ticks in Poland. Medical and Veterinary Entomology 28: 465-469. PDF

Bogdziewicz M., and R. Zwolak. 2014. Responses of small mammals to clear-cutting in temperate and boreal forests of Europe: a meta-analysis and review. European Journal of Forest Research 133: 1-11. PDF


Wróbel A., and R. Zwolak. 2013. The choice of seed tracking method influenced fate of beech seeds dispersed by rodents. Plant Ecology 214: 471-475. PDF

Zwolak R., S. Meagher, J. W. Vaughn, S. Dziemian, and E. E. Crone. 2013. Reduced ectoparasite loads of deer mice in burned forest: From fleas to trees? Ecosphere 4: art132. PDF


Zwolak R., and E. E. Crone. 2012. Quantifying the outcome of plant-granivore interactions. Oikos 121: 20-27. PDF

Zwolak R., D. E. Pearson, Y. K. Ortega, and E. E. Crone. 2012. Mechanisms driving postfire abundance of a generalist mammal. Canadian Journal of Zoology 90: 51-60. PDF


Zwolak R., D. E. Pearson, Y. K. Ortega, and E. E. Crone. 2010. Fire and mice: Seed predation reduces fire’s influence on conifer recruitment. Ecology 91: 1124-1131. PDF


Zwolak R. 2009. A meta-analysis of the effects of wildfire, clearcutting, and partial harvest on the abundance of North American small mammals. Forest Ecology and Management 258: 539-545. PDF

Zwolak R., and K. R. Foresman. 2008. Deer mouse demography in burned and unburned forest: no evidence for source-sink dynamics. Canadian Journal of Zoology 86: 83–91. PDF

Zwolak R., and K. R. Foresman. 2007. Effects of a stand-replacement fire on small mammal communities in montane forest. Canadian Journal of Zoology 85: 815-822. PDF

Zwolak R. 2007. Ogniu, krocz za mną: ekologia i polityka pożarow lasów Ameryki Północnej. Wiadomości Ekologiczne 53: 191–200.

Rychlik L., and R. Zwolak 2006. Interspecific behavioural domination among four sympatric species of shrews. Canadian Journal of Zoology 84: 434–448. PDF

Rychlik L., and R. Zwolak 2005. Behavioural mechanisms of conflict avoidance among shrews. Acta Theriologica 50: 289–308. PDF

Zwolak R., and L. Rychlik 2004. Does the reduction of locomotor activity serve as an aggression avoidance mechanism in shrews (Soricidae)? Electronic Journal of Polish Agricultural Universities 7(2), issue Biology.

Zwolak R. 2002. Zachowania agonistyczne ryjowkowatych - problem postaw ciała. Wiadomości Ekologiczne 48: 3–17.

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